BIOMIMETIC
MANUFACTURING OF FIBERS
INTRODUCTION
Biotechnology
provides the tools to clone and express designed synthetic protein fibers
in simple organisms. Spider dragline silk is a natural protein polymer
with many desirable properties, which we, and others, have used as a
model. Two partial cDNA sequences corresponding to the genes encoding
for the ß-sheet proteins (spidroin 1 and spidroin 2) of the dragline
silk of the golden orb weaver spider (Nephila clavipes) have been isolated
and published by other workers 1,2. Synthetic genes corresponding to
these natural genes were cloned, then expressed in E. coli 3,4, and
in yeast 5,6. A partial cDNA clone corresponding to the 3' end of the
dragline silk gene has also been isolated, cloned and expressed in E.
coli 7. Our goal is to produce silk-like material from spidroin 1 or
spidroin 2 homopolymers, and from spidroin 1/spidroin 2 or collagen/spidroin
heteropolymers by genetic engineering in yeast (Pichia pastoris) and
other plants (in particular, seed storage protein-producing plants where
it may be possible to produce significant quantities of these proteins).
We are using parts of the omega-helical collagen protein (mi-col-2 gene)
in the root-knot nematode Meloidogyne incognita cuticle8. For production
in plants, we are exploring the use of natural high level seed specific
promoters such as the omega-9 desaturase gene. The omega-9 desaturase
transcript has been shown to accumulate to very high concentrations
in peanut9 and sunflower10 developing seeds. We are now characterizing
genomic clones of this gene in peanut to obtain its promoter. A study
of the structural and functional biology of the spinning apparatus of
N. clavipes has been undertaken. While much of the study has been a
compilation of extant literature, we have conducted some studies in
our laboratory. The results of this study have been critical to the
design of our biomimetic spinning system. In addition, live spiders
are maintained in a walk-in enclosure and are “milked” for their dragline
silk. The current objective for study of the silk samples is to determine
the piezoelectric properties of the silk. We are also conducting Raman
spectroscopy and X-Ray Diffraction studies on the samples. The results
of this aspect of our program will support our goal to improve understanding
of the process/structure/property relations in natural spider silk,
and (eventually) in Biomimetic materials